Which of the following is a single-stranded DNA oligonucleotide that would move most slowly down an alkaline agarose gel during electrophoresis? 1) A short oligonucleotide with a high GC content 2) A long oligonucleotide with a high GC content 3) A short oligonucleotide with a high AT content 4) A long oligonucleotide with a high AT content